U-Pb ages of detrital zircons in Late Cretaceous sandstones overlying Paleozoic basement in southernmost Zealandia have unusually high proportions (40%) of Precambrian zircons, mostly Mesoproterozoic (ca. 1100. Ma) and Neoproterozoic (ca. 800. Ma). The proportion of Precambrian (Rodinia), relative to early Paleozoic (Gondwana), zircon groups is anomalously high and the former cannot have been reworked from any known basement rocks within Zealandia. Since Late Cretaceous sandstones were deposited after the breakup of Gondwana, it is proposed that their Precambrian zircons originated in Precambrian basement within Zealandia, briefly exposed along its Australian/Antarctic margin during Gondwana breakup. The ca. 1100. Ma and ca. 800. Ma age components suggest a former juxtaposition against the South China Block, when Zealandia was located east of Australia in Meso- and Neoproterozoic time.